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Full-grown, these monkeys are living proof of limbic regulation’s enduring power: they are timid, clingy, subordinate, and clumsy in their efforts to establish ties to other monkeys. The brains of these animals evidence permanent alterations in neurochemistry. Just because their mothers once lived under a pall of uncertainty, these adult animals show lifelong changes in levels of neurotransmitters like serotonin and dopamine. With their vulnerability to anxiety and depression, their social awkwardness and failures to attach as adults, these monkeys exhibit a close animal counterpart to the multifaceted misery that in human beings is labeled neurotic.